| Date | 1885 |
| Author | Loeffler |
| Contribution | With Frosch, found cause of foot-and-mouth disease. |
| Reference | Lechevalier |
| Notes | As with diphtheria, found weakness in Koch's postulates. In this case the problem was inability to demonstrate the organism (because it passed through filters!). Note date is approx. because not exact in L&S. |
| Category | Causation |
| Pathogen Class | General |
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