| Date | 1890 |
| Author | Winogradsky |
| Contribution | Isolated nitrifying bacteria from soil, confirming and extending work of others in a very significant way. |
| Reference | Doetsch |
| Notes | Others had failed to isolate such organisms because they assumed, erroneously, that they would thrive in the sort of highly organic media being used for the culture of animal pathogens. See 1877, 1881, 1889. |
| Category | Miscellany |
| Pathogen Class | Bacteria |
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