| Date | 1727 |
| Author | Hales |
| Contribution | Suggested transmission of hop mildew by small seeds. |
| Reference | Ainsworth |
| Notes | The Rev. Stephen Hales suggested both environmental causation and transmission by small seeds of mold. He considered it probable that seeds were blown by wind from the hops to the ground, thus making the ground a source of disease for subsequent hop crops. |
| Category | Causation |
| Pathogen Class | Fungi |
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