| Date | 1868 |
| Author | Keber |
| Contribution | Demonstrated the infectivity of vaccinia lymph after filtration, and observed objects of extremely small size. |
| Reference | Anon. |
| Notes | This is a very early isolation of a virus in the modern sense, but its significance could not be appreciated in its own time. The filter was "Swedish" filter paper. |
| Category | Causation |
| Pathogen Class | Viruses |
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