| Date | 1767 |
| Author | Targioni-Tozzetti |
| Contribution | Reported microscopic plant as cause of rust in wheat. |
| Reference | Ainsworth |
| Notes | Detailed work on black rust revealed microparasite in wheat plant. Concluded that transmission was by wind-borne seeds too small to be seen even by microscope. |
| Category | Causation |
| Pathogen Class | Fungi |
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