| Date | 1879 |
| Author | Klebs |
| Contribution | Inoculated syphilis into apes and probably saw the spirochaete. |
| Reference | Norman |
| Notes | T.A.E. Klebs. Published 1878-79. Schaudinn and Hoffmann reported the spirochaete in 1905. |
| Category | Causation |
| Pathogen Class | Bacteria |
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