| Date | 1895 |
| Author | Bruce |
| Contribution | Reported the tsetse fly as vector of the trypanosome that causes nagana in horses. |
| Reference | Foster |
| Notes | The role of tsetse fly in transmitting human trypanosomiasis was not demonstrated until the beginning of the 20th century. |
| Category | Causation |
| Pathogen Class | Bacteria |
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