| Date | 1886 |
| Author | Buist |
| Contribution | Reported that he had seen and measured viral elementary bodies in lymph taken from a cowpox inoculation ulcer. |
| Reference | Williams |
| Notes | George Brown Buist. Apparently the first observation of a virus in the modern sense; see 1868. The information was given in an oral presentation in 1886 and may have been published in 1887. |
| Category | Causation |
| Pathogen Class | Viruses |
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